All ASP.NET applications require at least one `Startup` class. When an application starts, ASP.NET searches the primary assembly for a class named `Startup` (in any namespace). You can specify a different assembly to search using the *Hosting:Application* configuration key. The class doesn't have to be `public`. If there are multiple `Startup` classes, ASP.NET looks for one in the project's root namespace, otherwise it chooses one in the alphabetically first namespace.
You can define separate `Startup` classes for different environments, and the appropriate one will be selected at runtime. Learn more in [Working with multiple environments](environments.md#startup-conventions).
The `Startup` class constructor can accept dependencies that are provided through [dependency injection](dependency-injection.md). You can use `IHostingEnvironment` to set up [configuration](configuration.md) sources and `ILoggerFactory` to set up [logging](logging.md) providers.
The `Startup` class must include a `Configure` method and can optionally include a `ConfigureServices` method, both of which are called when the application starts. The class can also include [environment-specific versions of these methods](environments.md#startup-conventions).
The `Configure` method is used to specify how the ASP.NET application will respond to HTTP requests. The request pipeline is configured by adding [middleware](middleware.md) components to an `IApplicationBuilder` instance that is provided by dependency injection.
In the following example from the default web site template, several extension methods are used to configure the pipeline with support for [BrowserLink](http://vswebessentials.com/features/browserlink), error pages, static files, ASP.NET MVC, and Identity.
Each `Use` extension method adds a [middleware](middleware.md) component to the request pipeline. For instance, the `UseMvc` extension method adds the [routing](routing.md) middleware to the request pipeline and configures [MVC](../mvc/index.md) as the default handler.
For more information about how to use `IApplicationBuilder`, see [Middleware](middleware.md).
Additional services, like `IHostingEnvironment` and `ILoggerFactory` may also be specified in the method signature, in which case these services will be [injected](dependency-injection.md) if they are available.
The `Startup` class can include a `ConfigureServices` method that takes an `IServiceCollection` parameter and optionally returns an `IServiceProvider`. The `ConfigureServices` method is called before `Configure`, as some features must be added before they can be wired up to the request pipeline.
For features that require substantial setup there are `Add[Something]` extension methods on `IServiceCollection`. This example from the default web site template configures the app to use services for Entity Framework, Identity, and MVC:
ASP.NET Core dependency injection provides application services during an application's startup. You can request these services by including the appropriate interface as a parameter on your `Startup` class's constructor or one of its `Configure` or `ConfigureServices` methods.